Jonathan, believed to be the world’s oldest known tortoise, died at an estimated age of 193 on the remote South Atlantic island of St Helena. The giant tortoise lived most of his life at Plantation ...

San Diego’s “Queen of the Zoo” has died. Her name was Gramma, and she was a Galápagos tortoise whose 141 years of existence (give or take) included more than 20 presidents, two world wars and a pair ...

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Corrections & clarifications: A previous version of this story incorrectly reported that Jonathan the tortoise had died. Despite claims of his death spreading on social media, the world's oldest known ...

SAN DIEGO — One of the first animals brought to the San Diego Zoo has died at the age of 141 years old. [ Read more trending news ] The zoo announced the death of its beloved Galápagos tortoise, ...

Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, has died on Nov. 20. She was (about) 141. The zoo's care team "made the compassionate and exceptionally ...

Gramma was born in her native habitat and was estimated to be about 141 years old, zoo officials said. She died Nov. 20. AP LOS ANGELES — After more than a century of munching on her favorite foods of ...